% =.Y BUL.LTIN 50.3 A * , r..;3 I AGRICULTUR AL EXPEI\IENT ST1ATIION/AUBURN UT\J\ LSITX AUBURN, ALABAMIA R. DENNIS ROUSE, IRECTOR CONTENTS Page EXPERIMENTAL METHODS RESULTS ............................................ Forage Yields ............................................... Botanical Composition of Forage ................................. Stand Persistence ..................... Parasitic Nematodes .................. ....................... ....................... ........... ....................... ............. 4 4 4 7 9 10 12 13 SUMMARY AND CONCLUSIONS ....................................... LITERATURE CITED ........................ ........................ FIRST PRINTING 4M, APRIL 1978 Information contained herein is available to everyone, regardless of race, color, or national origin. MAINTAINING FESCUE IN BAHIA MIXTURES C. S. HOVELAND, R. F. McCORMICK JR., E. L. CARDEN, R. RODRIGUEZ-KABANA, and J. T. SHELTON' PENSACOLA BAHIAGRASS (Paspalum notatum) is dormant and unproductive in Alabama from late October to April. Ken-. tucky 31 tall fescue (Festuca arundinacea)is semidormant in summer. It begins growth in September or October, grows little in mid-winter, and makes most of its production in spring until June or July. The two grasses in association should provide better forage over a longer grazing season than either one alone. Some farmers in central Alabama have successfully maintained the two species together in pastures over a number of years. More commonly, pure stands of tall fescue are eventually dominated by bahiagrass if it is introduced into the sward. Previous work in the southern Piedmont of Georgia (5) showed that tall fescue persisted and was productive in association with Coastal bermudagrass (Cynodon dactylon) for 3 years with high rates of nitrogen (N) fertilization. However, bahiagrass is generally considered to be more aggressive and forms a tighter sod than Coastal bermudagrass. Bahia is tolerant of close cutting or grazing in summer (4), while autumn production of tall fescue is reduced by close summer defoliation in Alabama (2). Tall fescue persistence and productivity is also reduced by nematodes on sandy soils (3). Thus, it is likely that management may be critical for maintenance of tall fescue in association with bahiagrass. This publication summarizes research in Alabama on the influence of rate and time of N application and stubble height on persistence and productivity of a tall fescue-bahiagrass mixture. Research Associate, Department of Agronomy and Soils; Superintendent, Gulf Coast Substation (formerly Superintendent, Brewton Experiment Field); Professor, Department of Botany and Microbiology; and Research Associate, Research Data Analysis. 'Professor, ALABAMA AGRICULTURAL EXPERIMENT STATION EXPERIMENTAL METHODS Pensacola bahiagrass seed were broadcast in March 1972 on Kentucky 31 tall fescue established in October 1971. The two experiments were located on Cahaba fine sandy loam (typic hapludult, fine-loamy siliceous) at the Plant Breeding Unit, Tallassee and on Benndale sandy loam (typic hapludult coarse-loamy siliceous) at the Brewton Experiment Field. During the years 1972-73, 1973-74, and 1974-75, nitrogen rates in pounds N per acre of 200 in winter and 200 in summer, 200 in winter and 100 in summer, 200 in winter and 0 in summer, 100 in winter and 100 in summer, 100 in winter and 0 in summer were applied in combination with summer (MaySept.) stubble heights of 4 and 11/2 inches, table 1. In addition, 200 and 100 pounds N per acre were applied in winter to summer rest plots (not cut from May through September). During the remainder of the year (October-April), all treatments were harvested to a stubble height of 11/2 inches. At the end of the summer, summer rest plots were cut and the residue, mostly dry tall fescue stems and leaves, was discarded. Winter N was applied in early October, early February, and late March. Summer N was applied in May, early July, and August. The experimental design was a randomized complete block, replicated four times. The percent tall fescue in forage was determined at each harvest. Percent tall fescue stand was estimated in April of the third year. Soil nematode populations were determined during the third year. RESULTS Forage Yields Winter-spring (October-April) forage yields, mainly tall fescue, were higher at the Plant Breeding Unit than at Brewton Experiment Field, tables 1 and 2. Yields during this season declined sharply over the 3-year period at both locations when plots were cut at the 1/2-inch stubble height. Yields also declined at lower rates of N, particularly at Brewton Experiment Field. Summer forage yields were highest when cut at the low stubble height during all 3 years of the tests. Results for 1975 illustrate effects of the various treatments on forage production, tables 1 and 2. Although summer rest increased winter-spring forage yield over no summer rest z -z z TABLE 1. SEASONAL DRY FORAGE PRODUCTION OF TALL FESCUE-BAHIAGRASS AS AFFECTED BY N FERTILIZATION AND STUBBLE HEIGHT OVER A 3-YEAR PERIOD AT THE PLANT BREEDING UNIT, TALLASSEE C con Summer stubble height Inches 4 N applied _________________________ Winter-spring yield 1973 T/acre 2.40 2.07 2.10 1.87 1.62 2.46 2.35 2.16 1.93 1.54 1.85 1.52 1974 Tiacre 1.91 1.73 1.57 0.81 0.61 1.82 1.40 1.08 0.86 0.59 1.81 1.05 1975 Tiacre 2.30 2.50 2.20 1.68 1.22 1.25 1.26 1.34 0.64 0.44 2.70 1.42 Winter Lb/acre 200 200 200 100 100 Summer Lb/acre 200 100 0 100 0 200 100 0 100 0 0 0 Summer yield ________________________ 1973 1974 1975 T/acre 2.01 2.10 0.44 1.12 0.70 4.41 3.25 2.14 3.41 2.05 0 0 T/acre 1.28 1.59 1.22 1.51 0.98 3.43 2.92 1.47 2.70 1.36 0 0 Tiacre 0.98 1.88 1.75 1.83 1.38 4.61 4.28 3.12 3.13 2.17 0 0 Total season yield c m 1973 Tiacre 4.46 3.17 2.58 2.99 2.32 6.87 5.60 4.30 5.34 3.64 1.34 1.02 1974 Tiacre 3.16 3.32 2.74 2.32 1.60 5.20 4.33 2.04 3.51 1.95 1.81 1.05 1975 Tiacre 3.28 4.36 3.87 3.02 2.60 5.92 5.53 4.41 3.78 2.62 2.70 1.42 con 11/2 200 200 200 100 100 Summer rest 200 100 icl TABLE 2. SEASONAL DRY FORAGE PRODUCTION OF TALL FESCUE-BAHIAGRASS AS AFFECTED BY N FERTILIZATION AND STUBBLE HEIGHT OVER A 3-YEAR PERIOD AT BREWTON EXPERIMENT FIELD Summer stubble height Inches 4 N applied Winter Summer Lb /acre Lb /acre 200 200 200 100 100 200 100 0 100 0 200 100 0 100 0 0 0 Winter-spring yield 1973 1974 1975 T/acre T/acre T/acre 1.53 1.32 1.14 1.15 0.72 0.82 0.80 0.66 0.55 0.46 1.43 1.02 1.12 1.00 0.78 0.46 0.23 0.64 0.63 0.50 0.32 0.16 1.18 0.34 0.46 0.53 0.60 0.22 0.22 0.22 0.14 0.27 0.08 0.07 0.98 0.30 Summer yield 1973 1974 1975 T/acre T/acre T/acre 2.88 2.17 0.64 2.04 0.71 4.40 3.22 1.38 3.17 1.21 0 0 4.92 4.16 2.25 3.66 1.64 6.88 5.27 3.21 4.58 2.09 0 0 5.96 5.34 3.57 4.17 2.22 6.71 5.75 4.28 4.69 2.68 0 0 Total season production 1973 1974 1975 T/acre T/acre T/acre 4.40 3.49 1.79 3.20 1.43 5.22 4.02 2.04 3.72 1.67 1.43 1.02 6.03 5.16 3.03 4.12 1.88 7.52 5.90 3.72 4.90 2.25 1.18 0.34 6.42 5.88 4.18 4.38 2.44 6.92 5.93 4.55 4.78 2.76 0.98 0.30 Ir 0 c C c r m 11/2 200 200 200 100 100 Summer rest 200 100 m m -I C,) -i -! a z 0 z MAINTAINING FESCUE IN BAHIA MIXTURES 7 (P<.01), it resulted in a large loss in potential summer production at both locations. Winter-spring forage yields were higher from plots cut in summer at a 4-inch as compared to a 1/2-inch stubble height. In contrast, bahiagrass production in summer was less when cut at the higher summer stubble height. This is because bahiagrass has such a large amount of forage growth in the 11 /2-to 4-inch zone. Total forage yields were highest when bahiagrass dominated the mixture, tables 1, 2, and 3. Highest average yield for the 3 years, 61/2 tons per acre, was at Brewton Experiment Field with summer stubble height of 11/2 inches and 400 pounds N per acre. Under this management at both locations, tall fescue was virtually eliminated by the second year, leaving nearly a solid stand of bahiagrass. Although higher winter N rates increased winter-spring production in the third year (P<.01), summer N had no effect on forage yields during this season, indicating that little N was carried over to the tall fescue in winter from the bahiagrass in summer. Summer-applied N rates resulted in a straight line increase in summer forage production of bahiagrass at both locations (P<.01). Reduced summer yields at the Plant Breeding Unit from the highest N rate and high stubble was a result sanguinalis) which was disof increased crabgrass (Digitaria carded when plots were harvested. This treatment resulted in little or no bahiagrass invasion but did permit crabgrass seedlings to become established in summer. Seasonal productivity during the third season (1975) with a tall fescue-bahia mixture was best with 200 pounds N per acre in winter and a summer stubble height of 4 inches, figure 1. This management system provided considerable winter and spring forage from tall fescue but resulted in reduced midsummer production. In contrast, cutting to 11/2 inches in summer resulted in low winter and spring production of tall fescue but high summer production from bahiagrass. Botanical Composition of Forage Tall fescue content of the total forage in each of the 3 years at the Plant Breeding Unit was highest with the 4-inch stubble height and 200 pounds N per acre in winter, table 3. Low stubble height drastically reduced the amount of tall fescue in forage, regardless of the N rate. After the first year at Brewton Experiment Field, tall fescue comprised only a small amount TABLE 3. PERCENT TALL FESCUE STAND IN 1975 AND PERCENTAGE TALL FESCUE IN TOTAL FORAGE AS AFFECTED BY N FERTILIZATION AND SUMMER STUBBLE HEIGHT OVER A 3-YEAR PERIOD AT PLANT BREEDING UNIT (PBU) AND BREWTON EXPERIMENT FIELD Summer stubble height Napplied Tall fescue stand in early April of 3rd year PBU Brewton 1973 Tall fescue composition of total forage 1974 1975 1973 1974 1975 a- Winter Summer Inches 4 Lb/acre 200 200 200 100 100 Lb/acre 200 100 0 100 0 200 100 0 100 0 0 0 % 89 90 89 76 35 23 46 39 11 6 100 58 % 19 38 40 20 35 0 4 14 5 6 62 45 %% 44 51 69 % 87 65 60 52 58 37 32 42 24 30 92 61 r 20 20 22 3 9 2 3 90 80 75 60 57 35 24 31 32 16 100 94 35 35 62 34 47 13 17 30 12 25 98 98 5 7 14 3 9 0 0 a 56 60 26 29 32 26 43 100 100 C I- 11/2 200 200 200 100 100 7 0 .0 88 56 3 0 0 89 83 r -o C Summer rest 200 100 I -o 0 z MAINTAINING FESCUE IN BAHIA MIXTURES Pounds per acre Tall fescue ] Bahiagrass HL HL HL HL HL HL HL Dec.18 Mar.26 Apr.24 June13 July 15 Aug12 Sept.30 FIG. 1. Seasonal forage distribution of tall fescue-bahia mixtures as af, fected by summer stubble heights of 4 inches (H) and 11/2 inches fertilized with 200 pounds N per acre in winter, and none in summer, third year, Plant Breeding Unit. of the forage, regardless of stubble height or N rate. Summer rest at both locations resulted in mainly tall fescue in the forage over the 3-year period. Stand Persistence Tall fescue stands in the central Alabama test remained excellent with high stubble height and high N fertilization, table 3. In contrast, stand and vigor of tall fescue was much less with low stubble height, figure 2. When N was applied at 100 pounds N per acre in winter-100 in summer or 100 in winter-none in summer, tall fescue stands were very poor. Summer rest maintained an excellent stand of tall fescue at the 200 pounds N per acre rate, figure 3, but the stand was much reduced at the 100 pounds N per acre rate. Tall fescue survival was much poorer at Brewton Experiment Field but treatment effects were similar to the other location. ALABAMA AGRICULTURAL EXPERIMENT STATION C, t 4 ' tf J l ' I r .. 4,L FIG.________ 2.Ctigt tbl nsme *2ic rgt eue tn n vigor oftl ecea:o prd octigi (let) [hnbt u mra er rw n soitonwt itr 4 -c stbl aiars n friie 5 95 wih20punsNprarei PatBedigUiMrc Fd.u.ruting tothir/2.ino he stu in sumr(ihrecedsn ,tble inisent a and nch stubbl mioreo tall fescue ast coredlto cuttin (let) whenu boh riei onemssatons wth bhgreadoferiliaed wpit 200 ponshe actreendiner Plnt Brieedn Ut, Marho 25,1975 singn oFtirld seiason. coilnhu iittr)an tn (Tycncorhtnc11Pareaitic weratpresetilags u- (Ne1magthes thrc lier o mjrtri the stLift .Iee\a tal~l fro ul indcaton ofhighe nominatce popsuationsr ai plrod containerng yeare. Tall feseu reuts veprunot coset. Lneatoes retrictindots to sha) nelat osoi rist rith preen uiiz at inentofiwaeda sprae depicth/(s Bairssts (3). andea to be hlmmortlerantl of) nematodes There unrdrotin lares n- MAINTAINING FESCUE IN BAHIA MIXTURES A.' ~~ ~ _________________ ~ ~ ~ ~ « d *'4 ,I, 1$ 6K k° FI.3 umrrs ftl _ -rq4 ecefetlzdwt 0 onsNprar tIG 3. Summer restbl tallfcu fertilized with pounds N per acre 200 (ih)we both were grown in association with bahiagrass. Plant Breeding Unit, March 25, 1975, spring of third season. N FritHil r n~ IN.t\n l B~i~i:s~ \i \ I liED MI oiI\ \ [Ii Illil Pi, }3tt BU (Pil) Ivin Si \inii i S "itt1111 Bii% lX io Suni I height r a pplicdl W\in~t(r Summiner B3cw01( Spring Atumnm ANv >\uiunnm Lh acr Lh (I acr) ~ o. Ni)o 100) 1'2 1) 41 24 65 2 10)0 100) 100( 0 (0 0) 1:3 12 31 44 52 24 6:3 Sumner est °_0 51 12 ALABAMA AGRICULTURAL EXPERIMENT STATION sandy soils, bahiagrass will have a competitive advantage and tall fescue will be eliminated. Management that favors bahiagrass, such as cutting or grazing to a low stubble in summer, will add to stress already present from nematode damage to tall fescue root systems. In an environment such as Brewton Experiment Field in southern Alabama with a long warm season combined with drought periods in spring and autumn, nematodes may easily weaken tall fescue plants and allow bahiagrass to dominate the sward. SUMMARY AND CONCLUSIONS 1. Experiments were conducted at the Plant Breeding Unit in central Alabama and at Brewton Experiment Field in southern Alabama over a 3-year period to determine the effect of N fertilizer rates and summer stubble heights on forage yield and persistence of tall fescue overseeded with bahiagrass. 2. Results of these experiments emphasize the difficulty of maintaining tall fescue stands in association with bahiagrass on sandy loam soils of central Alabama. Under high rates of winter N fertilization and a 4-inch summer stubble height, it was possible to maintain tall fescue with bahiagrass. This mixture provided forage over a longer season of the year than either species alone. Close clipping to a 11/2-inch stubble height in summer, together with low N fertilization rates, quickly resulted in complete bahiagrass dominance. 3. In southern Alabama on a sandy soil, tall fescue was replaced by bahiagrass within 2 years except with summer rest (not clipped May through September). 4. The high nematode populations suggest that nematodes may be a major reason for the demise of tall fescue in the mixture when drought stress or close summer defoliation favors bahiagrass. MAINTAINING FESCUE IN BAHIA MIXTURES LITERATURE CITED (1) 13 BEATY, E. R., R. L. STANLEY, AND J. POWELL. 1968. Effect of Height of Cut on Yield of Pensacola Bahiagrass. Agron. J. 60:356-358. (2) BERRY, R. F. AND C. S. HOVELAND. 1969. Summer Defoliation and Autumn-Winter Production of Phalaris Species and Tall Fescue Varieties. Agron. J. 61:493-497. (3) HOVELAND, C. S., R. RODRIGUEZ-KABANA, AND C. D. BERRY. 1975. Phalaris and Tall Fescue Forage Production as Affected by Nematodes in the Field. Agron. J. 67:714-717. (4) J. D. POWELL. 1967. Effect of Clipping Height on Forage Distribution and Regrowth of Pensacola Bahiagrass. Agron. J. 59:185-186. STANLEY, R. L., E. R. BEATY, AND (5) WILKINSON, S. R., L. F. WELCH, G. A. HILLSMAN, AND W. A. JACKSON. 1968. Compatibility of Tall Fescue and Coastal Bermudagrass as Affected by Nitrogen Fertilization and Height of Clip. Agron. J. 60:359-362. Alabama's Agricultural Experiment Station System AUBURN UNIVERSITY With an agricultural research unit in every major soil area, Auburn University serves the needs of field crop, livestock, forestry, and horticultural producers in each region in Alabama. Every citizen of the State has a stake in this research program, since any advantage from new and more economical ways of producing and handling farm products directly benefits the consuming public. O O 0 0 0 3 " s C 00 *Main 1 2 3 4 5 6 7 8. 9 10 11 Agricultural Experiment Station, Auburn. 12 13 14 15 16 17 18 19 20 Tennessee Valley Substation, Belle Mina. Sand Mountain Substation, Crossville North Alabama Horticulture Substation, Cullman Upper Coastal Plain Substation, Winfield Forestry Unit, Fayette County Thorsby Foundation Seed Stocks Farm, Thorsby Chilton Area Horticulture Substation, Clanton Forestry Unit, Coosa County Piedmont Substation, Camp Hill Plant Breeding Unit, Tallassee Forestry Unit, Autauga County Prattville Experiment Field, Prattville Black Belt Substation, Marion Junction Lower Coastal Plain Substation, Camden Forestry Unit, Barbour County Monroeville Experiment Field Monroeville Wiregrass Substation, Headland Brewton Experiment Field, Brewton Ornamental Horticulture Field Station. Spring Hill Gulf Coast Substation, Fairhope